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pulse perturba ti ons such as warm wate r large w ave or ma rine disease events modi fy the strength of local stabili zing feedba ck s thus predispo sing kelp forests to dispersal mediated synch rony of the ur chin ba rren stat e Kelp for ests a re sta bil ized through spor e producti on dis persal gro wth and the conversion t hrough senesc ence o f live kelp i nto drift algae which sea u rchins pr ef er to consum e and the refor e losi ng kelp i e losi ng the associated stabili zing feedbacks pa rticularly across large st retches of a coastl ine r educes the short term capa cit yofkelptorecove r Ifapertur bati onoccurssimul taneouslyortemporal lylags byacouple years a str ong s ea urch in recruit ment e vent th e combinati on can li mit kelp r ecov ery du e to conti nual grazer pressu re and subsequ ent dispersal mediat ed ur chin r ecruit ment can furt her stabi li ze the barren stat e Thus with a frame of syst ems th eory at it s core thi s project seeks to understand

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text:
how the localheterog eneit ystabili zingkelpforests int era ctswithpropaguleinput toprovideeit herr esil ient persist ence of the kelp fo rest state o r abrupt non li near movement tow ard s and a stabili zati on of the urchin ba rren st ate This int eracti on between local hete rogeneit y and propagule input is not st ati c but rather is anti cipated to be modified both by t he effects of keyston e predators such as seaott ers andbypulse perturbati onsandslow cha ngesinenvironmentalvariablesassociatedwith cli mate ch ange Ther efo re w e n eed to c apture c omm unit y information a cross a spati al extent overlappingare aswithandwithout seaott ers andweli kewiseneedtocaptu rethemannerinwhich these spati al patt erns re spond through ti me both to pulse perturbati ons and slowly changing environmental vari ables T h e p oten tial for sea ott ers to m itigate som e of t h e n egative ef fec ts of cli m ate ch an ge S ea ott ers exert profound trophic e ffe cts upon t emperate rocky re ef ecosyst ems via consum pti on ofinverteb ratep rey Inc ontrasttoall other marine mammals

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text:
thela ckofsig nificantadipos e ti ssue requires an extr emely high metaboli sm and caloric suppl y to balance the thermoregulatory costs ofli vingi natemperatemarineenvironment Thi stranslatestoavoracious appeti te andseaott ers are c apable o f consum ing 20 30 of their body weight a day with t he av erage Northern s ea ott er weighi ng 70 female to 90 male pounds Hale et al 2019 The indi rec t effects of such strong and direct consum er reso urce int e racti ons ripple t hroughout the benthos a nd in locati ons wher e sea ott ers a re the primary means of r egulating pop ulations of herbivorous s ea ur chins such as the Aleuti an Isl ands and pa rts of B rit ish C olum bia C anada sea ott er pr ese nce or absen ce is the differen ce betw een a div erse producti ve and hea lt hy kelp forest or a species depaupe rate state of urchin bar rens Estes Duggins 1995 Watso n Estes 2011 In other locati ons how ever particularly l owe r lati tude systems e g centr al C ali fornia the

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text:
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text:
cturall y we aken ed by increasing oce an acidi fi cati on and thus were more suscepti ble to sea urchin grazing And unfortunately a con com it ant warm wate r ev ent increas ed the r ate at whi ch sea ur chins graz ed upon the weakened alga e the net effe cts of the two physi cal variables chan neled through sea urchingrazingde cim ated the C lat hromorphum spp beds Theauthorsargu edthatifseaott ershad been pr esent to r egulate s ea ur chin abundan ce and behavior such a loss cou ld have be en avoid ed or significantl y mi ti gated Rasher et al 2020 A sim il ar theme is suggested b y the r ecent 20 13 anomalous warm wa ter event and subsequent nearshor e ec ologi cal fall out along stretches of the Northe ast P acific coast Despit e large sc alese astarloss alongtheWashing tonC oas tconcordantwiththeb ro aderS S WS event the benthos i e mobi leandsessi leinvertebratesandt urf understory andcano pymacroalgae withi n OCNMS seems to have fared bett e r in locati ons where se a ott ers ar e pres en t relative

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text:
sea ott ers consi stentl y forag es in proximi ty to S it e 5 and are consi ste ntl y absent from S it e 1 A healt hy b enthos at S it e 1 in 2011 with sessi le and mobi le invertebr ates and turf and und erstory a lgae B 2018 post S S WS the same ro ck at S it e 1scouredo falm ostall se ssi leinvertebrates andm acroalg aebys eaur chi ns C and D atS it e5wer e also t aken in 2011 and 2 018 respe cti vely and de spit e S S WS also occu rrin g the benthos along D retained the robust comm unit y of turf algae and se ssi le invertebrates 10 S ea ott er occupied c entr al C ali fornia is a test of the hypothesis regarding sea ott ers and resil ience and while k el p overall is doing b ett er there r elative to no rthern C ali fornia t he sp ati al scaleatwhichs eaott ers mit igatetheeffe ctsofurc hin barrensisnowinque sti on R ecentrese arc h demons trates se a ott ers a cti

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text:
vely avoid for aging on starving gonad l acking urchin bar ren u rchins and inst ead fo cus upon h ealt hy gravid u rchins wi thi n kelp forests Smi th et al in press Thus while there is sugg esti ve evidence that sea ott ers and pred ati on upon se a urchins in gen er al may regionally incr ease kelp forest r esil ience habit at specific foraging prefe renc es hav e the capa cit y to str engthen th e alre ady inhe rentl y patc hy dynami cs of k elp for ests at the loc al scal e Themannerinwhichthe sescale d epend entseaot terforaging eff ectsint era ctwithkelpforestand urchin bar ren f eedba ck processes such as kelp and urchin rec ruitm ent to s tructure comm unit y compos it ion i s enti rely unknown A new m od ali ty of b en thic m on itorin g to con fron t th e ch all en ges of a c h an gin g clim ate Here we propos e to esta bli sh a new par adigm of long term kelp for est moni toring

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text:
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text:
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