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ke lp fo rests where com bined densities o f the prostrate ke lps C tr iplicata H nigr ipes Costar ia and Agar um Neoagar um species ranged from almo st 18 ke lp per m² in the summ er to nearly 8 per m² in winter Juv enile kelps 30 cm in this study acco unte d fo r nearly all ke lp enco unte red in the winte r ranging fro m abo ut 6 t o 12 ke lp per m² In areas with a flo ating c ano py Ner eocyst is summ er densities o f nearly2 per m² were com plete ly absent during t hewint er Flo ating ke lp densities also v a ry seaso nally and can be lo we r than tho se of pro strate ke lps P eak Ner eocystis densities reco rded duringSCUBAsurv eys by Siddo net al 20 08 o f2individualsperm ²inthe 10 summ er and M axe ll and M iller 1 99 6 o f 2 indi v iduals per m ² in May were sim ilar to average surface densityo bserv atio nso f Nereocystis repo rte dby Callo way 2 01

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text:
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text:
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text:
rus pro ductio n o nce wat er tem perature s c o o l in late autum n Re dmo nd et al 20 14 Within 24 ho urs o f set tling spores germ inate into m ale and female gameto phy tes that m ature andsex ually re pro duceo v er the co urse o f o ne to two wee ks Steneck et al 200 2 Redmo nd et al 2 01 4 We rnberg et al 20 2 0 Once fertilized them icro sco pic juv enile spo ro phy te s growquickly becom ingv isibletothenakedeyeasafine fuzz afte r t hree to fo urweek s Redm o ndet al 2 01 4 M ax Calloway perso nal o bserv atio ns 12 Fig ur e 3 Diagram o f ke lp l ife stages sho w ing the juve nile an d adu lt s po ro phyte phas es a nd micro sc o pic z o o spo re and gameto phyte p hase s Ill ustratio n by L isa Sc harf Spit ler I n Mo n drago n J a nd J Mo ndra go n 20 03 Seaw

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text:
ee ds o f t he Pa c ific Co ast Co mmo n Marine Alga e fro m Alaska to B aja Califo rnia Sea C hallen ger s Mo nterey Cal ifo rnia 97 pages Ke lp fo rest seaso nal ity differ s depending o n species co m po sitio n and can ev en differ fro m po pulatio n to po pulatio no fthesamespecies Perennialspecies suchas P ter ygophora and giantke lp persistyear ro und while annualspecies like Costar ia appear anew e ach spring and disappear in t hewinter M att hews1 990 D ruehl and Clarksto n 20 00 Shaffer 20 00 A nnual species can som et im es persist lo nger and perennial species m ay fo llow annual cy cles In Washingto n ke lp po pulatio ns inside P uget So und and in the Strait o fGeo rgiaarelargely annual withperennialspecies including tho seo fthe S acchar ina com plex fo llo wing the annual cy cles o f Ner eocyst is and Costar ia M att hew s 1 99 0 M axe ll and M iller 19 94 In the Strait o f Juan de Fuca

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text:
Nereocystis cano pieso ften persist y ear ro und Shaffer2 00 3 Form Species Name Taxonomic synonyms Common name WA State Waters Puget Sound v ı o Egregia menziesii feather boa kelp Y Y Á ı x x Ç v ı P o P v o Y N Á ı ı À Æ µ v v o ı l v o Y Y À P v v ı À u v ı o l ı o Y N Á ı À Æ ˆ x ı u Y N ı À Æ ˆ x ı o v ı o l Á o Y Y x ı ı ı Y Y x x x ı ı Á v P o Y Y Á ı µ x x Costaria costata À o Y Y Á ı µ x x o v µ u Y N Á ı À Æ ˆ x Y N x x x Y Y x x v v o Y Y Á ı v ı Á v v o u o v ı Laminaria u o v ı Y Y x x x Z v µ P o Y Y Á ı À ˆ x Y Y Á ı

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text:
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text:
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text:
re biom ass per m² than seagrass T eagle et al 20 17 The to tal vo lum e o f habitat create d by a kelp fo rest is influenced by species m akeup ke lp density substrate av ailability and seaso n Maxe ll and M iller 19 96 Siddo n et al 2 00 8 Shaffer 2 000 Ke lp fo rests are no t nece ssarily higher quality habitat than ee lgrass o r salt m arshes no r is the relatio nship bet wee n these different nearsho re habitats antago nistic Sheaves et al 2 01 5 Instead kelp fo rest s are o ften o ne p iece o f larger scale habitat m o saics made up of patches o f ke lp fo rests eelgrass m eado ws and saltm arshes all o f who se seascape co nnect ivity increase s nearshore habitat quality and functio n as critical nurser ies Olso n et al 20 19 This is im po rtant to co nsider as kelp and ee lgrass require oppo site substrate s with ee lgrass requiring so ft m ud o r sand in which to lay

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text:
ro ots and kelps largely rely ing on hard substrate s including artificial structures bedro ck bo ulders gravel and co bble D espite these d ifferences in habitat requirements sev en o f nine surv ey respo ndents aswellastheautho r hav e enco unte red m ixedhabitats o f kelp and eelgrass Ke lps are characterized by rapid growt h m aking kelp forest s mo re pro ductiv e than rainfo rests or industrialagriculturefields K rum hanslet al 20 16 Thestructureprov idedby kelp serv esas a po te ntially edible habitat fo r inverte brates whose densities can be tw o to fiv e tim es greater in ke lp fo rests than in seagrass and o ther no n kelp seawe ed habitats Siddo n et al 200 8 Christie et al 20 09 Additio nally a single ke lp m ay pro v ide three distinct m icro habitats the ho ldfast stipe and blade that can each a tt ract 15 unique inv erte brate po pulatio ns Te agle et al 20 17 In Alaska increase d inv ertebrate abundances are asso ciate d with pro strate ke lp ev en when occurring belo

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