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inv o lv ed directly o r indirectly w ith HPA relate d activ ities Nine o f t he1 1 co ntacte d respo ndedto the survey and included sixWD FW st aff including t hree regio nal habitat bio lo gists two co nsultants Jen Jay Inc andM arine Surv eys Assessments M SA and o ne co unty planner P ierceCo unty Four o f t he nine respo ndentsrepo rted wo rki ng in all geographic regio ns and five inP uget So und two CP S o ne SP S o ne Whidbey Basin and o ne in CP S Hood Canal and t he Strait The o ute rco ast was not included as a cho ice in t he geo graphic regio n po rtio no fthe s urvey The surv ey quest io ns aimed to underst and t he degree o fknowledge o f no n flo ating k elp specie s distributio nsand assess po tential im pacts and asso ciate d degree s o fcertainty o r uncertainty o f SOWS related st ress effec ts to Washingto nke lpfo rests See Appendix B WD FW staff county planner

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text:
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text:
et al 2 002 We rnberg et al 2 02 0 The ov erall nativ e seaw eed dive rsity in the P acific No rthwe st is som e o f the highest in the wo rld and includes 22 species o f ke lp Gabrielso n et al 2 0 12 see Callo way et al 20 20 a for a det ailed list o f all 22 spe c ies Ke lp are bro wn m acro algae class P haeo phy ceae o f the o rder L aminariales that fo rm critical nearshore habitat fo r 6 num ero us prote cte d and endangered species in W ashingto n wate rs Calloway et al 2 020 b Hurd et al 2 01 4 All ke lp specie s co nsist o f a singl e o r m ultiple blades lam ina and a single o r m ultiple stipe stalk sec ured to hard substrate s by a ho ldfast By rely ing o n comm o n mo rpho lo gical features kelps can be o rganized into three broadcate go ries floating st alked stipitate and pro strate P ro strate

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text:
specie s generally co nsist o f a single wide blade and a single short generally less than appro ximate ly 15 cm stipe Stalke d species co nsist of single o r m ultiple blades held abov e the bentho s by a single o r m ultiple rigid wo o dy stipe s Flo ati ng specie s hav e m ultiple blades attached to single o r m ultiple stipes held aflo at by gas filled buo y s pneum ato cy sts O ther mo rpholo gical feat ures such as ho ldfast and blade structure can be used fo r species identificatio n and in o ur area are mo st he lpful for m o rpho lo gically sim ilar pro strate species fo r species identificatio n see D ruehl and Clarksto n 20 00 and Gabrielso n 20 12 fo r a more detailed taxo nom ic key Ho ldfasts are m ade up o f individual tangled branches hapte ra that resem ble ty pical plan t roo ts o r a single suctio n cup like disc and serv e o nly to att ach t he kelp to an

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text:
av ailable substrate P ro strate also called no n flo ating alo ng with stalked species kelps are by far the m o st specie s diverse abundant and widely distribute d in W ashingt on stat e wat ers T able 1 Table 2 In the expert inte rv iews andsurv ey co nductedfo rthisdo cum ent allrespo ndentsrepo rtedenco unte ringo neo rm ultiplepro strate species in HP A pro jec t areas regardless o f geo graphy Of pro strate species the m o st abundant recorded intheliteratureandrepo rtedby ex pertsandsurv ey respon dents areke lp s o fthe Sacchar ina Hedophyllum com plex hereafte r S acchar ina co m plex named fo r sugar ke lp S latissima fo rmerly L aminar ia sacchar ina a commo n kelp species fo und acro ss tem perate m arine areas thro ugho ut the no rthern hem isphere In addit io n to sugar ke lp S latiss ima t his co m plex includes S complanat a and H nigr ipes all o f which are genetically sim ilar and difficult if no t im po ssible to differentiate in the field Gabrielso n

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text:
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text:
iew The rem aining two flo ating species M acr ocystis and Ner eocystis fo rm m ixed flo ating surface cano pies alo ngtheo uterco astandtheStraito fJuandeFuca M o sto fo urunderst andingo fkelpbio lo gy ingeneral and ke lp fo rest eco lo gy co me s from research o n the large perennial M acr ocyst is fo rests alo ng the Califo rniac o ast InsideP uget So und Ner eocystis istheo nlyflo atingcano py fo rm ingspecies andhasbee n the fo cus o f recent c o nservatio n and research prio ritizatio n since the publishing of the P uget So und Ke lp Co nserv atio n and Recov ery P lan in 20 20 P uget So und Ner eocystis fo rest s have sustained do cumented lo sses in both so uth and c entral P uget So und Berry e t al 20 21 Berry unpublished 8 Table 1 Fro m W A DNR nd Sum mary o f Sho reZo ne inve nto ry 2000 submer ged a quatic veg etatio n data by c o unty Habitats dom inated by ke lp species are commo nly referred to

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text:
as kelp fo rest s o r ke lp beds depending o n u Z o P Ç u µ v ˆ v Z v u u v ı ı À l o ı _ v o Ç fo r tho se habitats with cano pies o f st alked o r floating species while po pulatio ns o f pro strate specie s are ˆ l o ˆ _ v v ˆ o Æ ı î ì í õ ˆ P v ı v µ v ˆ ı Ç _ u u v fo r both stipitate species gro wing under flo ating cano pies and fo r pro strate specie s in general These designatio ns do no t co nsider the fact that all kelp create s structure that pro v ide s refuge and abundant fo rage to associated species shade s the bentho s and require s sim ilar env iro nmenta l co nditio ns T eagle et al 2 01 7 We rnberget al 2 01 9 Thisdo cum entwillrefe rto allkelphabitatsaskelpfo rest s butpro v ide desc riptive distinctio ns when nece ssary e g no n floating ke lp fo rest

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text:
pro strate ke lp fo rest ke lp fo rest do m inated by S latiss ima et c There is no agreed upo n minim um density o f ke lps for a giv en po pulatio n to be co nsidered a fo rest o r a bed As We rnberg and Filbee Dex ter 2 02 0 po int o ut dete rm ining ex actly when a gro up o f individual seaw eedsbecom esam arine fo rest wo uldbeakinto askingho wm any co ralsittake sto m akeaco ralree f o rthe num bero fo yste rs required to fo rm ano yste rbed X _ Instead t he autho rsstre ss t hat a k elp fo rest is 9 any co llect io n o f ke lp species that fo rm a distinct subcano py env iro nme nt floating o r o therwise that pro v ides shelter and fo rage o ppo rtunities fo r asso ciate d species no t fo und in no n v eget ated habitats Whilethere are no co ncreteke lpdensity crite riafo rwhatm ake sake lpfo rest lim ite ddatafrom o urregio n can prov

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text:
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text:
m ultiple substrates and did no t target kelp po pulatio ns spec ifically At Titlow Beach in SPS M axe ll and M iller 199 6 tracked individual specie s densities o f Ner eocystis S latiss ima Costar ia and L aminar ia ephemer a fro m February 1 99 3 to M ay 19 94 Ke lp excluding the no n nativ e no n ke lp S muticum density peak edinJune1 9 9 6 atappro xim ately 9 kelp individuals per m² andwasreducedto 0 5 ke lp individuals per m² by D ecem ber 1 99 6 M axell and M iller 19 95 Figure 2 S latissim a formerly L sacchar ina wasby farthem ostabundantke lpspeciesprese nt reaching peakdensitieso fappro xim ately 5 5individualsper m² inJune19 96 M axellandM iller19 96 Figure 2 Cost ar ia theseco ndmo stabundant species ov er the study perio d attained densities o f clo se to 2 individuals per m² Ner eocystis density in M ay wasalso c loseto 2 individuals perm ² Siddo n et al 2 00 8 o bserved sim ilar seaso nal trends but higher densities o f prostrate ke lps in Alaska n

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